Proposal to conserve the name Juenllia, nom. cons., against an additional name Urbania (Verbenaceae)

O'LEARY, NATALY; PERALTA, PAOLA; MÚLGURA DE ROMERO, MARÍA E.

TAXON

INT ASSOC PLANT TAXONOMY

Lugar: Viena; Año: 2009 vol. 58 p. 655 - 655

0040-0262

Urbania was established as a new genus in Verben aceaeby Philippi in 1891 (in honor of the German botanist I. Urban),based in two species: U. pappigera and U. eganioides. It wasaccepted as such by Reiche (in Fl. Chile 5. 1910) and Troncoso(in Darwiniana 18: 295?412. 1974). Botta (in Bol. Soc.Argent. Bot. 25: 477?483. 1988) synonymised U. eganioidesunder U. pappigera rendering Urbania a monotypic genusconfined to the high Andean Puna of Argentina and Chile.Moldenke (l.c.) published the name Junellia (in honorof the German botanist S. Junell) with 35 species previouslyincluded in Verbena, and this genus has since been widelyrecognised in many South American floristic publications,taxonomic revisions and cytogenetic studies such as Covas& Schnack in Revista Argent. Agron. 13: 153?166. 1946, inHaumania 1: 32?41. 1947; Cabrera in Dimitri, Encicl. Argent.Agric. Jard. 2(1): 1?85. 1976; Raj in Rev. Paleobot. Palynol.39: 361. 1983; Beck in Ecol. Bolivia 6: 8. 1985; Poggio & al.in Bol. Soc. Argent. Bot. 25: 542?545. 1988; Botta in Darwiniana29: 373?377. 1989, in Cabrera, Flora Jujuy 13(9):63?76. 1993, in Correa, Flora Patagónica 8(6): 147?195. 1999;Botta & Brandham in Kew Bull. 48: 143?150. 1993; Martínez& al. in Darwiniana 34: 1?17. 1996; Atkins in Kubitzki, Fam.Gen. Vasc. Pl. 7: 449?468. 2004, and Peralta & al. in Ann.Missouri Bot. Gard. 95: 338?390. 2008.Currently the name Junellia applies to a genus of 39species and 6 varieties (Peralta & al., l.c.) that grow in SouthAmerica from Perú and Bolivia to Argentina and Chile. Thehistory of the name was briefly reviewed by Botta & al. (inTaxon 44: 639?640. 1995) in a proposal to conserve Junelliaagainst Monopyrena Speg. and Thryothamnus Phil., whichhas since been recommended (Brummitt in Taxon 47: 863?872. 1998) and accepted and included first in the St. LouisCode (Greuter & al. in Regnum Veg. 138. 2000).Junellia differs from Urbania only by the presence oflong hygroscopic hairs in the calyx of the latter. Phylogeneticstudies using morphological characters (Peralta & al.in Resumenes V Reunión Argentina Cladística: 48. 2004)show that the monotypic genus Urbania falls within a cladecomprising species of Junellia. Recent molecular phylogeneticstudies, being carried on by O?Leary & al. (in Syst.Bot., in press), based on four cloroplast regions (trn L-F, trnS-G, trn S-fm, trn D-T) and the nuclear ITS and ETS, confirmthe nested position of Urbania within a monophyleticclade of Junellia. Hence the name Urbania threatens thename Junellia.Urbania is itself also conserved (McNeill & al. in RegnumVeg. 146: 410. 2006) against the earlier Urbania Valtke(in Oesterr. Bot. Z. 25: 10. 1875). Conservation of Junelliaover Urbania would not have any impact on this other conservationand would permit the continued use of Urbaniaby anyone wishing to recognise it as distinct from Junellia.It is here proposed to conserve the name JunelliaMoldenke, considering the wide use of this name, therebyavoiding the need for 39 new combinations.