Cladistic analysis of the South American species of Tricorythodes (Ephemeroptera: Leptohyphidae) with descriptions of new species and stages

MOLINERI, C.

AQUATIC INSECTS

Año: 2002 vol. 24 p. 273 - 308

0165-0424

Fifteen species of the genus Tricorythodes are treated: T. arequita Traver, T. australis (Banks), T. barbus Allen, T. bullus Allen, T. cristatus Allen, T. curiosus (Lugo-Ortiz & McCafferty) comb. nov., T. hiemalis Molineri, T. mirca sp. nov., T. nicholsae (Wang et al.) comb. nov., T. ocellus Allen & Roback, T. popayanicus Domínguez, T. quizeri sp. nov., T. santarita Traver, T. yura sp. nov. and T. zunigae sp. nov. A matrix of 17 taxa and 28 characters was analized cladistically using the program PIWE (Parsimony with Implied Weights, Goloboff 1993a). Haplohyphes furtiva Domínguez and Leptohyphes eximius Eaton were used to root the tree. Only one tree of maximum fit was obtained supporting the new combinations T. curiosus and T. nicholsae, the monophily of the genus, and the invalidity, at least in part, of Allen & Murvosh´s (1987) subgenera. The following species are described for the first time from all the stages: T. mirca sp. nov., T. quizeri sp. nov., T. yura sp. nov. and T. zunigae sp. nov. The following stages are described for the first time: nymphs and imagines of both sexes of T. arequita, nymph and female imago of T. santarita, male and female imagines of T. bullus. The following stages are redescribed: nymphs of T. barbus and female imago of T. santarita. Distributional records, diagnoses and keys to distinguish all the South American species of Tricorythodes are given.