New Paleogene Marsupials from the Amazonian Basin of Southeastern Perú

GOIN AND CANDELA

The Paleogene Mammalian

Natural History Museum of Los Angeles County

Año: 2004; p. 15 - 60

An analysis of 79 fossil specimens, most comprising tiny, isolated upper and lower molars recovered from Paleogene levels near Santa Rosa in the Peruvian Amazon, led to the recognition of eight new genera and eleven new species of extinct marsupials: Rumiodon inti gen. et sp. nov. (Didelphimorphia, cf. Herpetotheriidae), Patene campbelli sp. nov. (Sparassodonta, Hathliacynidae), Incadolops ucayali gen. et sp. nov. (Polydolopimorphia, Prepidolopidae), Wamradolops tsullodon gen. et sp. nov. (Polydolopimorphia, family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), nov. (Polydolopimorphia, Prepidolopidae), Wamradolops tsullodon gen. et sp. nov. (Polydolopimorphia, family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), campbelli sp. nov. (Sparassodonta, Hathliacynidae), Incadolops ucayali gen. et sp. nov. (Polydolopimorphia, Prepidolopidae), Wamradolops tsullodon gen. et sp. nov. (Polydolopimorphia, family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), nov. (Polydolopimorphia, Prepidolopidae), Wamradolops tsullodon gen. et sp. nov. (Polydolopimorphia, family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Rumiodon inti gen. et sp. nov. (Didelphimorphia, cf. Herpetotheriidae), Patene campbelli sp. nov. (Sparassodonta, Hathliacynidae), Incadolops ucayali gen. et sp. nov. (Polydolopimorphia, Prepidolopidae), Wamradolops tsullodon gen. et sp. nov. (Polydolopimorphia, family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), nov. (Polydolopimorphia, Prepidolopidae), Wamradolops tsullodon gen. et sp. nov. (Polydolopimorphia, family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), sp. nov. (Sparassodonta, Hathliacynidae), Incadolops ucayali gen. et sp. nov. (Polydolopimorphia, Prepidolopidae), Wamradolops tsullodon gen. et sp. nov. (Polydolopimorphia, family indeterminate), Hondonadia pittmanae sp. nov. (Polydolopimorphia, family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), family indeterminate), Perulestes cardichi and P. fraileyi gen. et spp. nov. (Paucituberculata, Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), Caenolestidae), Sasawatsu mahaynaq gen. et sp. nov. (Paucituberculata, cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae), cf. Palaeothentidae), Kirutherium paititiensis gen. et sp. nov. (Microbiotheria, Microbiotheriidae),